Milkcaps and Brittlegills
The family Russulaceae contains two genera: Lactarius and Russula. In Europe there are some 160 species in each of these genera. The two groups differ mainly in that Lactarius fungi produce a milky latex from their gills when damaged whereas Russula species do not. Both genera are characterised by brittle flesh and pale spores.
Russula and Lactarius species are different from most other mushrooms because rather than consisting of tube-like cells they contain spherical cells called sphaerocysts, making their texture less pliable and more fragile than other mushrooms.
Russula mushrooms - the Brittlegills The Russula genus contains mushrooms of many colours, from white, yellow, tan and brown, through pink, orange red and purple to blue, mauve and even green. All brittlegills are ectomycorrhizal with trees or shrubs, and so they occur only in woodland settings or with shrubby plants (including, in some instances, heathers and other heathland species). The species illustrated here is Russula xerampelina; its cap colours are very variable - a characteristic of many of the brittlegills.
Brittlegills are fleshy fungi. Most of them are edible, although many species are too hot and acrid for most people's taste, and some - Russula nobilis for example - cause stomach upsets.
Lactarius mushrooms - the Milkcaps The Lactarius mushrooms are known as milk caps because of the latex that is exuded from their gills when they are cut or broken. Like the brittlegills, milkcaps are mycorrhizal fungi, and they occur with trees or shrubs.
Illustrated here are Oakbug Milkcaps, Lactarius quietus, which is a very common species found in oak woodland.
The initial colour of the milk, its taste, and any colour change as the milk dries are valuable identification characters in a group with little cap colour variation.
Russula fungi are sometimes referred to as russules rather than brittlegills, while milkcaps are in older field guides generally referred to as milk caps or milk-caps.